13 PALEOBIOS, VOL 21, NUMBER 2, SEPTEMBER 2001 Fig. 1. Dorsal (left) and ventral (or palmar; right) views of I.ACM 144725. Scale bar 1 either arthritis (an infectious process) or arthrosis (a non- infectious process). The pathogenesis of non-articular periostoses of this type in equids has been discussed by Rooncy (1997). In general these lesions are related to avulsion of ligaments from their bony attachments as a result of instability and jerky, irregu- lar movements of the affected leg or leg segment (Rooncy and Robertson 1996). Such instability usually results from misplacement of the foot on a surface. ACKNOWLEDGEMENTS The authors thank Samuel A. Mcl-cod and Gary T. Takcuchi of the Natural History Museum of Los Angeles County, who provided LACM 144725. Thanks are also ex- tended to volunteers Alex and Matthew Van Dam of the Natural History Museum of Los Angeles County, who or- ganized the collection of fossil horses from San Josccito Cave and discovered the specimen. LITERATURE CITED Azzaroli, A. 1995. A synopsis of the Quaternary species of Equus in North America. Bolletino delta Societa Pakontohgica Italians 34(2):205-221. Azzaroli, A. 1998. The genus Equus in North America: The Pleis- tocene species. Paleontojiraphica halica 85:1-60. Bennett, D.K. 1980. Stripes do not a zebra make. Part I -A cla- distic analysis of Equus. Systematic Zoology 29:272 287. Dalquest, W.W. 1978. Phylogeny of American horses of Blancan and Pleistocene age. AnnatesZookgici Fennici 15(3): 191 199. Kurten, B., and E. Anderson. 1980. Pleistocene mammals of North America. Columbia University Press, New York. 442 pp. Rooney, J.R. 1997. Equid paleopathology. Journal of Equine Vet- erinary Science 17(8):430-446. Rooney, J.R. 1999. Veterinary paleopathology, pp. 392-394 in McGraw-Hill Yearbook of Science and Technology. McGraw- Hill, New York. Rooney, J.R., and J.I.. Robertson. 1996. Equine Pathology. Iowa State University Press, Ames, Iowa. 502 pp. Rooney, JR., and E. Scott. 1998. Sidebone in a fossil horse. Jour- nal of Equine Veterinary Science 18:519 520. Scott, E. 1996. The small horse from Valley Wells, San Bernar- dino County, California. San Bernardino County Museum As- sociation Quarterly 43( l,2):85-89. Scott, E. 1997. A review of Equus conversidens in southern Cali- fornia, with a report on a second, previously unrecognized species of Pleistocene small horse from the Mojave Desert. SCOT!' & ROON^r-PRRIOSTOSLS IN EQUUS PHALANX 14 Fig. 2. Radiograph of LACM 144725 in dorsal view. The negative image (original radiograph) is on the left; positive on the right. See text for description of features. Journal of Vertebrate Paleontology 17(3):75A. Stock, C. 1950. 25,000 year old horse: The skeleton ofan Ice Age horse makes a return trip to Mexico. Etiijinccrinjj and Science Monthly 14:16-17. Winans, M.C. 1989. A quantitative study of North American fos- sil species of the genus Y.quus. pp. 262 297 in D.R. Prothero and R.M. Schoch (eds.). The Evolution of Perissodactyls. Oxford University Press, Oxford. Zietzschmann, O., E. Ackernecht, and II. Cirau. 1943. Ellenberger Baum. I Iandbuch der Vergleichenden Anatomic Der I laustiere, 18'1' ed. Springcr-Verlag, Berlin.